*136850
FUMARATE HYDRATASE; FH

Alternative titles; symbols
FUMARASE

Other entities represented in this entry:
FUMARATE HYDRATASE, CYTOSOLIC, INCLUDED; FH1, INCLUDED
FUMARATE HYDRATASE, MITOCHONDRIAL, INCLUDED; FH2, INCLUDED

HGNC Approved Gene Symbol: FH

Cytogenetic location: 1q43     Genomic coordinates (GRCh37): 1:241,660,856 - 241,683,084 (from NCBI)

Gene Phenotype Relationships
Location Phenotype Phenotype
MIM number
1q43 Fumarase deficiency 606812
Leiomyomatosis and renal cell cancer 150800

TEXT
Description
Fumarate hydratase, or fumarase (EC 4.2.1.2), is an enzymatic component of the tricarboxylic acid, or Krebs, cycle. It catalyzes the conversion of fumarate to malate.

Enzyme Structure

Fumarase exists in both cytosolic and mitochondrial forms, which differ in electrophoretic mobility. Kinsella and Doonan (1986) found an unusually high degree of identity of structure of human fumarase and that from B. subtilis and E. coli. Using peptide mapping, O'Hare and Doonan (1985) showed that the cytosolic and mitochondrial fumarases from pig liver are identical over nearly all of their amino acid sequences, but that they differ at their N-termini.

Isoenzymes

Edwards and Hopkinson (1979) studied a family with an electrophoretic variant of FH. Two persons had variation in both the soluble and the mitochondrial forms, suggesting that they are determined by a single locus. Posttranslational processing may account for the differences between the 2 forms. Another possibility is that the 2 enzymes are heteromeric and share a subunit coded by chromosome 1. If posttranslational processing is the mechanism, all control of processing (e.g., enzymes) seems to be coded by chromosome 1. Common genetic control of both soluble and mitochondrial fumarate hydratase is further supported by the finding of combined deficiency. Doonan et al. (1984) cited evidence suggesting that the isoenzymes of fumarase are translated in precursor form from 2 different mRNA molecules, these mRNAs in turn arising from alternative splicing of a single gene transcript.

Mapping
Van Someren et al. (1974) and Craig et al. (1976) found that the fumarase locus is on chromosome 1, possibly in the area 1q42. Despoisses et al. (1984) narrowed the regional assignment of FH to 1q42.1 by gene dosage studies in patients with various types of partial trisomy or partial monosomy of 1q. Coughlin et al. (1993) mapped the FH gene to chromosome 1 using PCR-amplified cDNA as a probe in Southern blots of genomic DNA from a series of mouse/human somatic cell hybrids. They observed related sequences on chromosomes 13 and 5.

Gene Function
Pollard et al. (2005) stated that the nuclear-encoded Krebs cycle enzymes fumarate hydratase and succinate dehydrogenases (see, e.g., SDHB 185470) act as tumor suppressors, and germline mutations in these genes predispose individuals to leiomyomas and renal cancer and to paragangliomas (see 115310), respectively. Pollard et al. (2005) showed that FH-deficient cells and tumors accumulated fumarate and, to a lesser extent, succinate. SDH-deficient tumors principally accumulated succinate. In situ analysis showed that these tumors also overexpressed HIF1A (603348), activation of HIF1A targets like VEGF (192240), and high microvessel density. Pollard et al. (2005) hypothesized that increased succinate and/or fumarate may stabilize HIF1A, and that the basic mechanism of tumorigenesis in paragangliomas and leiomyoma and renal cancer may be pseudohypoxic drive, just as it is in von Hippel-Lindau syndrome (193300).

Using genetically modified mouse kidney cells in which Fh1 had been deleted, Frezza et al. (2011) applied a newly developed computer model of the metabolism of these cells to predict and experimentally validate a linear metabolic pathway beginning with glutamine uptake and ending with bilirubin excretion from Fh1-deficient cells. This pathway, which involves the biosynthesis and degradation of heme, enables Fh1-deficient cells to use accumulated tricarboxylic acid (TCA) cycle metabolites and permits partial mitochondrial NADH production. Frezza et al. (2011) predicted and confirmed that targeting this pathway would render Fh1-deficient cells nonviable, while sparing wildtype Fh1-containing cells. Frezza et al. (2011) concluded that their work went beyond identifying a metabolic pathway that is induced in Fh1-deficient cells to demonstrate that inhibition of heme oxygenation is synthetically lethal when combined with Fh1 deficiency, providing a potential target for treating HLRCC (150800) patients.

Molecular Genetics
In patients with fumarase deficiency (606812), Bourgeron et al. (1993) and Coughlin et al. (1993) identified mutations in the FH gene (136850.0001 and 136850.0002).

In patients with multiple cutaneous and uterine leiomyomata (150800), Tomlinson et al. (2002) identified mutations in the FH gene (136850.0003 and 136850.0004).

In patients with hereditary leiomyomatosis and renal cell cancer, Tomlinson et al. (2002) identified several mutations in the FH gene (136850.0005 and 136850.0006).

Using sequence analysis, Toro et al. (2003) identified germline mutations in the FH gene in 31 of 35 (89%) families with cutaneous leiomyomas. Eighteen of the 20 different mutations they identified-- 2 insertions, 5 small deletions that caused frameshifts leading to premature truncation of the protein, and 13 missense--were novel. The same mutation, arg190 to his (R190H; 136850.0007), was identified in 11 unrelated families. Cutaneous leiomyomas were found in 81 individuals (47 women and 34 men). Uterine leiomyomas were also found in 98% (46 of 47) of women with cutaneous leiomyomas. Total hysterectomy was performed in 89% (41 of 46) of women with cutaneous and uterine leiomyomas, 44% before or at age 30 years. In 13 individuals in 5 families, Toro et al. (2003) identified unilateral and solitary renal tumors. Papillary type II renal cell carcinoma was present in 7 individuals from 4 families, and another individual from 1 of these families had collecting duct carcinoma of the kidney. The study expanded the histologic spectrum of renal tumors and FH mutations associated with hereditary leiomyomatosis and renal cell carcinoma.

Barker et al. (2002) analyzed a series of 26 leiomyosarcomas and 129 uterine leiomyomas (from 21 patients) for somatic mutations in fumarate hydratase and allelic imbalance around 1q43. None of the 26 leiomyosarcomas harbored somatic mutations in fumarate hydratase. Only 5% (7 of 129) of the leiomyomas showed allele imbalance at 1q42-q43, and no somatic mutations in fumarate hydratase were observed.

Alam et al. (2003) reported 20 FH mutations in 35 of 46 probands with multiple cutaneous and uterine leiomyomata (MCUL) or FH deficiency. Disease-associated missense FH changes mapped to highly conserved residues, mostly in or around the enzyme's active site or activation site. The mutation spectra in FH deficiency and MCUL were similar, although in the latter mutations tended to occur more 5-prime in the gene and were predicted to result in a truncated or absent protein. The authors reported that not all mutation-carrier parents of FH deficiency children had a strong predisposition to leiomyomata. Renal carcinoma is sometimes part of MCUL, as part of the variant hereditary leiomyomatosis and renal cancer (HLRCC) syndrome; these cancers may have either type II papillary or collecting duct morphology. There was no association between the type or site of FH mutation and any aspect of the MCUL phenotype. Biochemical assay for reduced FH functional activity in the germline of MCUL patients may indicate carriers of FH mutations with high sensitivity and specificity, and can detect reduced FH activity in some patients without detectable FH mutations. The authors concluded that MCUL is probably a genetically homogeneous tumor predisposition syndrome, primarily resulting from absent or severely reduced fumarase activity.

To determine whether FH mutations may predispose women to developing nonsyndromic uterine leiomyomas (UL; 150669), Gross et al. (2004) performed a genetic linkage study with DNA from 123 families containing at least 1 affected sister pair. In addition, to assess the frequency of FH loss specifically in uterine leiomyomas with 1q rearrangements, they performed a FISH analysis of UL. Analysis of the genotyping data revealed evidence suggestive of linkage to the FH region among study participants who were less than 40 years of age at diagnosis (p = 0.04). FISH results showed that 1 copy of FH was absent in 9 of 11 ULs. Gross et al. (2004) concluded that loss of FH may be a significant event in the pathogenesis of a subset of nonsyndromic ULs.

Because some individuals with HLRCC with a germline FH mutation have breast cancer (114480), Kiuru et al. (2005) analyzed germline FH mutations from 85 Finnish breast cancer patients, most of whom were selected based on positive family or personal history for malignancies associated with HLRCC. No mutations were found. Kiuru et al. (2005) concluded that FH is not a major predisposing gene for familial breast cancer.

Wei et al. (2006) identified 14 mutations in the FH gene, including 9 novel mutations, in affected members of 13 families with HLRCC and 8 families with multiple cutaneous and uterine leiomyomata. Four unrelated families had the R58X mutation (136850.0003) and 5 unrelated families had the R190H mutation (136850.0007). Cutaneous leiomyomata were present in 16 (76%) of 21 families, ranging from mild to severe. All 22 female mutation carriers from 16 families had uterine fibroids. Renal tumors occurred in 13 (62%) of 21 families. No genotype/phenotype correlations were identified.

To examine the cancer risk and tumor spectrum in Finnish families positive for FH mutations, Lehtonen et al. (2006) collected genealogic and cancer data from 868 individuals. FH mutation status was analyzed in all 98 available patients. The standardized incidence ratio (SIR) was 6.5 for renal cell carcinoma (RCC) and 71 for uterine leiomyosarcoma (ULMS). The overall cancer risk was statistically significantly increased in the age group of 15 to 29 years, consistent with features of cancer predisposition families in general. An FH germline mutation was found in 55% of studied individuals. Most RCC and ULMS displayed biallelic inactivation of FH, as did breast and bladder cancers. In addition, Lehtonen et al. (2006) observed several benign tumors including atypical uterine leiomyomas, kidney cysts, and adrenal gland adenomas.

As part of the French National Cancer Institute study, Gardie et al. (2011) identified 32 different heterozygous germline mutations in the FH gene, including 21 novel mutations, in 40 (71.4%) of 56 families with proven HLRCC. In addition, FH mutations were found in 4 (17.4%) of 23 probands with isolated type 2 papillary renal cell carcinoma, including 2 patients with no family history. In vitro functional expression studies showed that all mutations caused about a 50% decrease in FH enzymatic activity. In addition, there were 5 asymptomatic mutation carriers in 3 families, indicating incomplete penetrance. The findings indicated that renal call carcinoma can be the only manifestation of this disorder. No genotype/phenotype correlations were identified.

ALLELIC VARIANTS (Selected Examples):

.0001 FUMARASE DEFICIENCY
FH, ALA265THR

In a patient of Arab ancestry with fumarase-deficiency (606812), Coughlin et al. (1993) identified a G-to-A transition at nucleotide 793 changing ala265 to thr (A265T). The father was shown to be heterozygous for the mutation.

.0002 FUMARASE DEFICIENCY
FH, GLU319GLN

Bourgeron et al. (1993, 1994) described a glu319-to-gln (E319Q) mutation in the FH gene in 2 daughters of first-cousin Moroccan parents who presented with progressive encephalopathy, dystonia, leukopenia, and neutropenia at an early age. Elevation of lactate in the cerebrospinal fluid (so-called hyperlactatorachia) and high fumarate excretion in the urine led Bourgeron et al. (1994) to investigate the activities of the respiratory chain and of the Krebs cycle, and finally to identify fumarase deficiency (606812). The deficiency was profound, was present in all tissues investigated, and affected the cytosolic and mitochondrial isoenzymes to the same degree. The sibs were homozygous for a missense mutation, a G-to-C transversion at nucleotide 955. The predicted amino acid substitution occurred in a highly conserved region of the fumarase cDNA. Both parents exhibited half the expected fumarase activity in their lymphocytes and were found to be heterozygous for the mutation.

.0003 HEREDITARY LEIOMYOMATOSIS AND RENAL CELL CANCER
FH, ARG58TER

In 3 families, Tomlinson et al. (2002) found that members affected by multiple cutaneous and uterine leiomyomata (150800) had a change of codon 58 from CGA (arg) to TGA (stop) (R58X) in exon 2 of the FH gene.

In 3 unrelated families with hereditary leiomyomatosis and renal cell cancer, Wei et al. (2006) identified the R58X mutation, resulting from a 172C-T transition. The R58X mutation was also identified in affected members of a fourth unrelated family with multiple cutaneous and uterine leiomyomata. Haplotype analysis of the families did not show a founder effect, suggesting that R58X represents a hot spot mutation.

.0004 HEREDITARY LEIOMYOMATOSIS AND RENAL CELL CANCER
FH, ASN64THR

In 6 separate families, Tomlinson et al. (2002) found that individuals with multiple cutaneous and uterine leiomyomata (150800) were heterozygous for a mutation in codon 64 in exon 2 of the FH gene converting AAC (asn) to ACC (thr) (N64T).

In a 55-year-old man with hereditary leiomyomatosis and renal cell cancer and the N64T mutation in the FH gene, Carvajal-Carmona et al. (2006) identified a Leydig cell tumor of the testis.

.0005 HEREDITARY LEIOMYOMATOSIS AND RENAL CELL CANCER
FH, 2-BP DEL

In 2 Finnish families with the hereditary leiomyomatosis and renal cell cancer syndrome (150800), Tomlinson et al. (2002) found a 2-bp deletion in codon 181 in exon 4 of the FH gene: conversion of GAGTTT to GTTT.

.0006 HEREDITARY LEIOMYOMATOSIS AND RENAL CELL CANCER
FH, ARG300TER

In a Finnish family with the hereditary leiomyomatosis and renal cell cancer syndrome (150800), Tomlinson et al. (2002) found a nonsense mutation converting codon 300 in exon 6 of the FH gene from CGA (arg) to TGA (stop) (R300X).

.0007 HEREDITARY LEIOMYOMATOSIS AND RENAL CELL CANCER
FH, ARG190HIS

In 4 individuals from a family with cutaneous and uterine leiomyomatosis and renal cell cancer (HLRCC; 150800), Toro et al. (2003) identified a 569G-A transition in exon 4 of the FH gene, resulting in an arg190-to-his (R190H) mutation. The R190H mutation was also present in 10 other unrelated families with cutaneous and uterine leiomyomatosis, but screening for occult renal tumors in affected individuals from these 10 families did not identify renal tumors. Thus there appeared to be other genetic and/or environmental factors that influenced the phenotype.

Wei et al. (2006) identified the R190H mutation in affected members of 3 unrelated families with HLRCC. The R190H mutation was also identified in affected members of 2 additional families with multiple cutaneous and uterine leiomyomata. A founder effect could not be determined.

.0008 HEREDITARY LEIOMYOMATOSIS AND RENAL CELL CANCER
FH, ARG190LEU

Toro et al. (2003) described a family with leiomyomatosis and renal cell cancer (150800) associated with a 569G-T transition in exon 4 of the FH gene, resulting in an arg190-to-leu (R190L) mutation. The nucleotide substitution occurred at the same position as that changed in the common R190H mutation (136850.0007).

.0009 HEREDITARY LEIOMYOMATOSIS AND RENAL CELL CANCER
FH, ARG58PRO [dbSNP:rs75086406]

In affected members of a family with multiple cutaneous and uterine leiomyomata (150800), Chan et al. (2005) identified a heterozygous 173G-C transversion in exon 3 of the FH gene, resulting in an arg58-to-pro (R58P) substitution. The proband was a 77-year-old Polish woman with multiple cutaneous leiomyomas and uterine fibroids. Her eldest daughter had a similar phenotype, and 2 unaffected daughters did not have the mutation. Her son had multiple skin leiomyomas and was diagnosed with metastatic papillary renal cell cancer at age 50 years, and his asymptomatic 20-year-old son was also found to carry the mutation and was thus likely to develop skin leiomyomas, but the risk of renal cancer was difficult to predict. Chan et al. (2005) noted that a nonsense mutation in the same residue had been reported (R58X; 136850.0003).

Heinritz et al. (2008) identified the R58P mutation in affected members of a large German family with multiple cutaneous and uterine leiomyomata without renal cancer. Family history revealed that this German family originally came from Poland but was dispersed after World War II. Haplotype analysis of this family and that reported by Chan et al. (2005) demonstrated a founder effect for the mutation.

See Also:
Busby et al. (1976); Edwards and Hopkinson (1979); Petrova-Benedict et al. (1987); Tolley and Craig (1975); van Someren et al. (1974)

REFERENCES
1. Alam, N. A., Rowan, A. J., Wortham, N. C., Pollard, P. J., Mitchell, M., Tyrer, J. P., Barclay, E., Calonje, E., Manek, S., Adams, S. J., Bowers, P. W., Burrows, N. P., and 18 others. Genetic and functional analyses of FH mutations in multiple cutaneous and uterine leiomyomatosis, hereditary leiomyomatosis and renal cancer, and fumarate hydratase deficiency. Hum. Molec. Genet. 12: 1241-1252, 2003. [PubMed: 12761039, related citations] [Full Text: HighWire Press, Pubget]

2. Barker, K. T., Bevan, S., Wang, R., Lu, Y.-J., Flanagan, A. M., Bridge, J. A., Fisher, C., Finlayson, C. J., Shipley, J., Houlston, R. S. Low frequency of somatic mutations in the FH/multiple cutaneous leiomyomatosis gene in sporadic leiomyosarcomas and uterine leiomyomas. Brit. J. Cancer 87: 446-448, 2002. [PubMed: 12177782, related citations] [Full Text: Nature Publishing Group, Pubget]

3. Bourgeron, T., Chretien, D., Poggi-Bach, J., Doonan, S., Rabier, D., Letouze, P., Munnich, A., Rotig, A., Landrieu, P., Rustin, P. Mutation of the fumarase gene in two siblings with progressive encephalopathy and fumarase deficiency. J. Clin. Invest. 93: 2514-2518, 1994. [PubMed: 8200987, related citations] [Full Text: Journal of Clinical Investigation, Pubget]

4. Bourgeron, T., Chretien, D., Rotig, A., Munnich, A., Landrieu, P., Rustin, P. Molecular characterization of fumarase deficiency in two children with progressive encephalopathy. (Abstract) Am. J. Hum. Genet. 53 (suppl.): A891 only, 1993.

5. Busby, N., Courval, J., Francke, U. Regional assignments of the genes for fumarate hydratase and guanylate kinase on chromosome 1 and for lysosomal acid phosphatase and esterase A4 on chromosome 11. Cytogenet. Cell Genet. 16: 105-107, 1976. [PubMed: 185008, related citations] [Full Text: Pubget]

6. Carvajal-Carmona, L. G., Alam, N. A., Pollard, P. J., Jones, A. M., Barclay, E., Wortham, N., Pignatelli, M., Freeman, A., Pomplun, S., Ellis, I., Poulsom, R., El-Bahrawy, M. A., Berney, D. M., Tomlinson, I. P. M. Adult Leydig cell tumors of the testis caused by germline fumarate hydratase mutations. J. Clin. Endocr. Metab. 91: 3071-3075, 2006. [PubMed: 16757530, related citations] [Full Text: HighWire Press, Pubget]

7. Chan, I., Wong, T., Martinez-Mir, A., Christiano, A. M., McGrath, J. A. Familial multiple cutaneous and uterine leiomyomas associated with papillary renal cell cancer. Clin. Exp. Derm. 30: 75-78, 2005. [PubMed: 15663510, related citations] [Full Text: Blackwell Publishing, Pubget]

8. Coughlin, E. M., Chalmers, R. A., Slaugenhaupt, S. A., Gusella, J. F., Shih, V. E., Ramesh, V. Identification of a molecular defect in a fumarase deficient patient and mapping of the fumarase gene. (Abstract) Am. J. Hum. Genet. 53 (suppl.): A896 only, 1993.

9. Craig, I., Tolley, E., Bobrow, M. Mitochondrial and cytoplasmic forms of fumarate hydratase assigned to chromosome 1. Cytogenet. Cell Genet. 16: 118-121, 1976. [PubMed: 975868, related citations] [Full Text: Pubget]

10. Despoisses, S., Noel, L., Choiset, A., Portnoi, M.-F., Turleau, C., Quack, B., Taillemite, J.-L., de Grouchy, J., Junien, C. Regional mapping of FH to band 1q42.1 by gene dosage studies. (Abstract) Cytogenet. Cell Genet. 37: 450-451, 1984.

11. Doonan, S., Barra, D., Bossa, F. Structural and genetic relationships between cytosolic and mitochondrial isoenzymes. Int. J. Biochem. 16: 1193-1199, 1984. [PubMed: 6397370, related citations] [Full Text: Pubget]

12. Edwards, Y. H., Hopkinson, D. A. Further characterization of the human fumarase variant, FH2-1. Ann. Hum. Genet. 43: 103-108, 1979. [PubMed: 525970, related citations] [Full Text: Pubget]

13. Edwards, Y. H., Hopkinson, D. A. The genetic determination of fumarase isozymes in human tissues. Ann. Hum. Genet. 42: 303-313, 1979. [PubMed: 434773, related citations] [Full Text: Pubget]

14. Frezza, C., Zheng, L., Folger, O., Rajagopalan, K. N., MacKenzie, E. D., Jerby, L., Micaroni, M., Chaneton, B., Adam, J., Hedley, A., Kalna, G., Tomlinson, I. P. M., Pollard, P. J., Watson, D. G., Deberardinis, R. J., Shlomi, T., Ruppin, E., Gottlieb, E. Haem oxygenase is synthetically lethal with the tumour suppressor fumarate hydratase. Nature 477: 225-228, 2011. [PubMed: 21849978, related citations] [Full Text: Nature Publishing Group, Pubget]

15. Gardie, B., Remenieras, A., Kattygnarath, D., Bombled, J., Lefevre, S., Perrier-Trudova, V., Rustin, P., Barrois, M., Slama, A., Avril, M.-F., Bessis, D., Caron, O., and 41 others. Novel FH mutations in families with hereditary leiomyomatosis and renal cell cancer (HLRCC) and patients with isolated type 2 papillary renal cell carcinoma. J. Med. Genet. 48: 226-234, 2011. Note: Erratum: J. Med. Genet. 48: 576 only, 2011. [PubMed: 21398687, related citations] [Full Text: HighWire Press, Pubget]

16. Gross, K. L., Panhuysen, C. I. M., Kleinman, M. S., Goldhammer, H., Jones, E. S., Nassery, N., Stewart, E. A., Morton, C. C. Involvement of fumarate hydratase in nonsyndromic uterine leiomyomas: genetic linkage analysis and FISH studies. Genes Chromosomes Cancer 41: 183-190, 2004. [PubMed: 15334541, related citations] [Full Text: John Wiley & Sons, Inc., Pubget]

17. Heinritz, W., Paasch, U., Sticherling, M., Wittekind, C., Simon, J. C., Froster, U. G., Renner, R. Evidence for a founder effect of the germline fumarate hydratase gene mutation R58P causing hereditary leiomyomatosis and renal cell cancer (HLRCC). Ann. Hum. Genet. 72: 35-40, 2008. [PubMed: 17908262, related citations] [Full Text: Blackwell Publishing, Pubget]

18. Kinsella, B. T., Doonan, S. Nucleotide sequence of a cDNA coding for mitochondrial fumarase from human liver. Biosci. Rep. 6: 921-929, 1986. [PubMed: 3828494, related citations] [Full Text: Pubget]

19. Kiuru, M., Lehtonen, R., Eerola, H., Aittomaki, K., Blomqvist, C., Nevanlinna, H., Aaltonen, L. A., Launonen, V. No germline FH mutations in familial breast cancer patients. Europ. J. Hum. Genet. 13: 506-509, 2005. [PubMed: 15523491, related citations] [Full Text: Nature Publishing Group, Pubget]

20. Lehtonen, H. J., Kiuru, M., Ylisaukko-oja, S. K., Salovaara, R., Herva, R., Koivisto, P. A., Vierimaa, O., Aittomaki, K., Pukkala, E., Launonen, V., Aaltonen, L. A. Increased risk of cancer in patients with fumarate hydratase germline mutation. J. Med. Genet. 43: 523-526, 2006. [PubMed: 16155190, related citations] [Full Text: HighWire Press, Pubget]

21. O'Hare, M. C., Doonan, S. Purification and structural comparisons of the cytosolic and mitochondrial isoenzymes of fumarase from pig liver. Biochim. Biophys. Acta 827: 127-134, 1985. [PubMed: 3967032, related citations] [Full Text: Pubget]

22. Petrova-Benedict, R., Robinson, B. H., Stacey, T. E., Mistry, J., Chalmers, R. A. Deficient fumarase activity in an infant with fumaricacidemia and its distribution between the different forms of the enzyme seen on isoelectric focusing. Am. J. Hum. Genet. 40: 257-266, 1987. [PubMed: 3578275, related citations] [Full Text: Pubget]

23. Pollard, P. J., Briere, J. J., Alam, N. A., Barwell, J., Barclay, E., Wortham, N. C., Hunt, T., Mitchell, M., Olpin, S., Moat, S. J., Hargreaves, I. P., Heales, S. J., and 9 others. Accumulation of Krebs cycle intermediates and over-expression of HIF1-alpha in tumours which result from germline FH and SDH mutations. Hum. Molec. Genet. 14: 2231-2239, 2005. [PubMed: 15987702, related citations] [Full Text: HighWire Press, Pubget]

24. Tolley, E., Craig, I. Presence of two forms of fumarase (fumarate hydratase EC 4.2.1.2) in mammalian cells: immunological characterisation and genetic analysis in somatic cell hybrids; confirmation of the assignment of a gene necessary for the enzyme expression to human chromosome 1. Biochem. Genet. 13: 867-883, 1975. [PubMed: 812482, related citations] [Full Text: Pubget]

25. Tomlinson, I. P. M., Alam, N. A., Rowan, A. J., Barclay, E., Jaeger, E. E. M., Kelsell, D., Leigh, I., Gorman, P., Lamlum, H., Rahman, S., Roylance, R. R., Olpin, S., and 19 others. Germline mutations in FH predispose to dominantly inherited uterine fibroids, skin leiomyomata and papillary renal cell cancer. Nature Genet. 30: 406-410, 2002. [PubMed: 11865300, related citations] [Full Text: Nature Publishing Group, Pubget]

26. Toro, J. R., Nickerson, M. L., Wei, M.-H., Warren, M. B., Glenn, G. M., Turner, M. L., Stewart, L., Duray, P., Tourre, O., Sharma, N., Choyke, P., Stratton, P., Merino, M., Walther, M. M., Linehan, W. M., Schmidt, L. S., Zbar, B. Mutations in the fumarate hydratase gene cause hereditary leiomyomatosis and renal cell cancer in families in North America. Am. J. Hum. Genet. 73: 95-106, 2003. [PubMed: 12772087, related citations] [Full Text: Elsevier Science, Pubget]

27. van Someren, H., Van Henegouwen, H. B., de Wit, J. Evidence for synteny between the human loci for fumarate hydratase, UDG glucose pyrophosphorylase, 6-phosphogluconate dehydrogenase, phosphoglucomutase-1, and peptidase-C in man-Chinese hamster somatic cell hybrids. Cytogenet. Cell Genet. 13: 150-152, 1974. [PubMed: 4827484, related citations] [Full Text: Pubget]

28. van Someren, H., Van Henegouwen, H. B., Westerveld, A., Bootsma, D. Synteny of the human loci for fumarate hydratase and UDPG pyrophosphorylase with chromosome 1 markers in somatic cell hybrids. Cytogenet. Cell Genet. 13: 551-557, 1974. [PubMed: 4549862, related citations] [Full Text: Pubget]

29. Wei, M.-H., Toure, O., Glenn, G. M., Pithukpakorn, M., Neckers, L., Stolle, C., Choyke, P., Grubb, R., Middelton, L., Turner, M. L., Walther, M. M., Merino, M. J., Zbar, B., Linehan, W. M., Toro, J. R. Novel mutations in FH and expansion of the spectrum of phenotypes expressed in families with hereditary leiomyomatosis and renal cell cancer. J. Med. Genet. 43: 18-27, 2006. [PubMed: 15937070, related citations] [Full Text: HighWire Press, Pubget]

Contributors: Ada Hamosh - updated : 9/21/2011
George E. Tiller - updated : 11/21/2008
Cassandra L. Kniffin - updated : 10/6/2008
John A. Phillips, III - updated : 6/21/2007
Victor A. McKusick - updated : 7/5/2006
Cassandra L. Kniffin - updated : 2/13/2006
Victor A. McKusick - updated : 4/26/2005
George E. Tiller - updated : 3/9/2005
Victor A. McKusick - updated : 12/20/2004
Victor A. McKusick - updated : 6/25/2003
Victor A. McKusick - updated : 10/23/2002
Cassandra L. Kniffin - reorganized : 4/4/2002
Victor A. McKusick - updated : 2/28/2002
Creation Date: Victor A. McKusick : 6/4/1986
Edit History: alopez : 09/22/2011
terry : 9/21/2011
ckniffin : 9/13/2011
wwang : 8/16/2011
ckniffin : 8/11/2011
wwang : 11/21/2008
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ckniffin : 10/6/2008
carol : 6/21/2007
alopez : 7/7/2006
terry : 7/5/2006
wwang : 2/28/2006
ckniffin : 2/13/2006
tkritzer : 4/29/2005
terry : 4/26/2005
alopez : 3/9/2005
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alopez : 10/24/2002
terry : 10/23/2002
alopez : 4/12/2002
ckniffin : 4/4/2002
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carol : 4/4/2002
ckniffin : 4/4/2002
ckniffin : 4/4/2002
terry : 3/27/2002
alopez : 3/1/2002
terry : 2/28/2002
terry : 7/24/1998
jason : 6/15/1994
terry : 4/27/1994
carol : 10/28/1993
carol : 10/18/1993
supermim : 3/16/1992
carol : 11/20/1990