Entry - *178642 - SURFACTANT, PULMONARY-ASSOCIATED PROTEIN A2; SFTPA2 - OMIM
 
* 178642

SURFACTANT, PULMONARY-ASSOCIATED PROTEIN A2; SFTPA2


Alternative titles; symbols

PULMONARY SURFACTANT PROTEIN AII; SPAII; SPA2
COLLECTIN 5; COLEC5


HGNC Approved Gene Symbol: SFTPA2

Cytogenetic location: 10q22.3   Genomic coordinates (GRCh38) : 10:79,555,852-79,560,407 (from NCBI)


Gene-Phenotype Relationships
Location Phenotype Phenotype
MIM number
Inheritance Phenotype
mapping key
10q22.3 Interstitial lung disease 2 178500 AD 3

TEXT

Cloning and Expression

Katyal et al. (1992) isolated and characterized SFTPA2, which encodes an isoform of surfactant-associated protein SPA (SFTPA1; 178630). SFTPA1 and SFTPA2 both contain a consensus recognition sequence for initiation, TATAAA, and a polyadenylation signal sequence. Transcripts of both genes were found in adult human lung by Northern blot analysis.

Wang et al. (2009) noted that the 248-amino acid SFTPA2 protein contains a collagen-like region, a neck domain, and a C-terminal carbohydrate-recognition domain. SFTPA2 shares greater than 98% amino acid identity with SFTPA1 across the coding region and greater than 90% amino acid identity across noncoding regions.


Gene Structure

Katyal et al. (1992) determined that the SFTPA2 gene contains 5 exons.


Mapping

Kolble et al. (1993) presented evidence for a cluster of pulmonary surfactant-associated proteins on chromosome 10q22-q23. They constructed a phylogenetic tree showing the relationships of SPAI (SFTPA1), SPAII, mannose-binding lectin (MBL; 154545), SPB (178640), and SPC (178620) proteins.

Stumpf (2021) mapped the SFTPA2 gene to chromosome 10q22.3 based on an alignment of the SFTPA2 sequence (GenBank BC111571) with the genomic sequence (GRCh38).


Molecular Genetics

Surfactant protein-A, which plays a role in innate host defense in the lung, is also expressed in the eustachian tube. Ramet et al. (2001) reported that the frequency of specific surfactant protein-A haplotypes and genotypes differs between children with recurrent otitis media compared with a control population in Finland.

In affected members of 2 families with interstitial lung disease-2 (ILD2; 178500), some of whom also had lung cancer, Wang et al. (2009) identified 2 heterozygous missense mutations in the SFTPA2 gene (G231V, 178642.0001 and F198S, 178642.0002, respectively). Both mutations involve highly conserved residues in the carbohydrate recognition domain (CRD) and were predicted to disrupt protein structure; transfection studies in A549 human alveolar epithelial cells showed that the mutant proteins were retained in the endoplasmic reticulum and not secreted.

In 11 probands with ILD2, Legendre et al. (2020) identified heterozygous missense mutations in the SFTPA2 gene (see, e.g, 178642.0003-178642.0005). The mutations, which were found by direct sequencing, were either absent from or found at a low frequency in the gnomAD database. The mutation segregated with the disorder in families from whom DNA was available; there was evidence of incomplete penetrance and variable expressivity. In vitro functional expression studies in HEK293T cells transfected with the mutations showed normal protein expression with decreased secretion of SFTPA1 compared to controls. Abnormal cytoplasmic retention of mutant SFTPA1 in the alveolar epithelium was considered to contribute to pathogenicity.


ALLELIC VARIANTS ( 5 Selected Examples):

.0001 INTERSTITIAL LUNG DISEASE 2

SFTPA2, GLY231VAL
  
RCV000014086...

In affected members of a 4-generation family (F27) with interstitial lung disease-2 (ILD2; 178500), some of whom also had lung cancer, Wang et al. (2009) identified heterozygosity for a c.692G-T transversion in exon 6 of the SFTPA2 gene, resulting in a gly231-to-val (G231V) substitution at a highly conserved residue in the carbohydrate recognition domain (CRD). The mutation was not found in 3,557 population-based controls. Transfection studies in A549 human alveolar epithelial cells showed that the mutant protein was retained in the endoplasmic reticulum and not secreted.


.0002 INTERSTITIAL LUNG DISEASE 2

SFTPA2, PHE198SER
  
RCV000014087

In a 45-year-old Caucasian male (CKG810) with interstitial lung disease-2 (ILD2; 178500) and lung cancer (see 211980), Wang et al. (2009) identified heterozygosity for a c.593T-C transition in exon 6 of the SFTPA2 gene, resulting in a phe198-to-ser (F198S) substitution at a highly conserved residue in the carbohydrate recognition domain (CRD). The proband reported multiple family members with pulmonary fibrosis and/or lung cancer, but they were not available for study. The patient had no personal smoking history, but smoking status of his childhood home was unclear. The mutation was not found in 3,557 population-based controls. Transfection studies in A549 human alveolar epithelial cells showed that the mutant protein was retained in the endoplasmic reticulum and not secreted.


.0003 INTERSTITIAL LUNG DISEASE 2

SFTPA2, ASN171ILE
  
RCV001780092

In a patient (family 4) with interstitial lung disease-2 (ILD2; 178500), Legendre et al. (2020) identified a heterozygous c.512A-T transversion (c.512A-T, NM_001098668.2) in exon 6 of the SFTPA2 gene, resulting in an asn171-to-ile (N171I) substitution in the CRD domain. The mutation, which was found by direct sequencing, was not found in the gnomAD database. In vitro functional expression studies in HEK293T cells transfected with the mutation showed normal protein expression with decreased secretion of SFTPA1 compared to controls. Abnormal cytoplasmic retention of mutant SFTPA1 in the alveolar epithelium was considered to contribute to pathogenicity. The patient had a significant family history of similar lung disease, but DNA from affected family members was not available for segregation studies.


.0004 INTERSTITIAL LUNG DISEASE 2

SFTPA2, VAL178MET (rs371035540)
  
RCV001265772...

In 3 patients from 2 unrelated families (families 5 and 6) with interstitial lung disease-2 (ILD2; 178500), Legendre et al. (2020) identified a heterozygous c.532G-A transition (c.532G-A, NM_001098668.2) in exon 6 of the SFTPA2 gene, resulting in a val178-to-met (V178M) substitution in the CRD domain. The mutation, which was found by direct sequencing, was found at a low frequency in the gnomAD database (3 of 250,974 alleles). One unaffected family member in family 5 carried the mutation, consistent with incomplete penetrance. In vitro functional expression studies in HEK293T cells transfected with the mutation showed normal protein expression with decreased secretion of SFTPA1 compared to controls. Abnormal cytoplasmic retention of mutant SFTPA1 in the alveolar epithelium was considered to contribute to pathogenicity. The proband in family 6 had 2 similarly affected deceased family members, including those with lung cancer, but DNA from those individuals was not available for segregation studies.


.0005 INTERSTITIAL LUNG DISEASE 2

SFTPA2, TRP233ARG
  
RCV001784080

In a mother and son (family 8) with interstitial lung disease-2 (ILD2; 178500), Legendre et al. (2020) identified a heterozygous c.697T-A transversion (c.697T-A, NM_001098668.2) in exon 6 of the SFTPA2 gene, resulting in a trp233-to-arg (W233R) substitution in the CRD domain. The mutation, which was found by direct sequencing, was not present in the gnomAD database. In vitro functional expression studies in HEK293T cells transfected with the mutation showed normal protein expression with decreased secretion of SFTPA2 compared to controls. Abnormal cytoplasmic retention of mutant SFTPA2 in the alveolar epithelium was considered to contribute to pathogenicity. There was a significant family history of similar lung disease, including lung cancer, but DNA from affected family members was not available for segregation studies.


REFERENCES

  1. Katyal, S. L., Singh, G., Locker, J. Characterization of a second human pulmonary surfactant-associated protein SP-A gene. Am. J. Resp. Cell Molec. Biol. 6: 446-452, 1992. [PubMed: 1372511, related citations] [Full Text]

  2. Kolble, K., Lu, J., Mole, S. E., Kaluz, S., Reid, K. B. M. Assignment of the human pulmonary surfactant protein D gene (SFTP4) to 10q22-q23 close to the surfactant protein A gene cluster. Genomics 17: 294-298, 1993. [PubMed: 8406480, related citations] [Full Text]

  3. Legendre, M., Butt, A., Borie, R., Debray, MP., Bouvry, D., Filhol-Blin, E., Desroziers, T., Nau, V., Copin, B., Dastot-Le Moal, F., Hery, M., Duquesnoy, P., and 29 others. Functional assessment and phenotypic heterogeneity of SFTPA1 and SFTPA2 mutations in interstitial lung diseases and lung cancer. Europ. Resp. J. 56: 2002806, 2020. [PubMed: 32855221, related citations] [Full Text]

  4. Ramet, M., Lofgren, J., Albo, O.-P., Hallman, M. Surfactant protein-A gene locus associated with recurrent otitis media. J. Pediat. 138: 266-268, 2001. [PubMed: 11174628, related citations] [Full Text]

  5. Stumpf, A. M. Personal Communication. Baltimore, Md. 11/16/2021.

  6. Wang, Y., Kuan, P. J., Xing, C., Cronkhite, J. T., Torres, F., Rosenblatt, R. L., DiMaio, J. M., Kinch, L. N., Grishin, N. V., Garcia, C. K. Genetic defects in surfactant protein A2 are associated with pulmonary fibrosis and lung cancer. Am. J. Hum. Genet. 84: 52-59, 2009. [PubMed: 19100526, images, related citations] [Full Text]


Anne M. Stumpf - updated : 11/18/2021
Cassandra L. Kniffin - updated : 11/15/2021
Marla J. F. O'Neill - updated : 2/19/2009
Ada Hamosh - updated : 4/23/2001
Creation Date:
Victor A. McKusick : 9/27/1993
carol : 11/19/2021
alopez : 11/18/2021
ckniffin : 11/15/2021
wwang : 02/27/2009
terry : 2/19/2009
mgross : 3/10/2003
cwells : 5/9/2001
terry : 4/23/2001
mcapotos : 6/8/2000
mcapotos : 6/7/2000
carol : 6/19/1998
terry : 6/18/1998
mimadm : 4/13/1994
carol : 9/27/1993

* 178642

SURFACTANT, PULMONARY-ASSOCIATED PROTEIN A2; SFTPA2


Alternative titles; symbols

PULMONARY SURFACTANT PROTEIN AII; SPAII; SPA2
COLLECTIN 5; COLEC5


HGNC Approved Gene Symbol: SFTPA2

Cytogenetic location: 10q22.3   Genomic coordinates (GRCh38) : 10:79,555,852-79,560,407 (from NCBI)


Gene-Phenotype Relationships

Location Phenotype Phenotype
MIM number
Inheritance Phenotype
mapping key
10q22.3 Interstitial lung disease 2 178500 Autosomal dominant 3

TEXT

Cloning and Expression

Katyal et al. (1992) isolated and characterized SFTPA2, which encodes an isoform of surfactant-associated protein SPA (SFTPA1; 178630). SFTPA1 and SFTPA2 both contain a consensus recognition sequence for initiation, TATAAA, and a polyadenylation signal sequence. Transcripts of both genes were found in adult human lung by Northern blot analysis.

Wang et al. (2009) noted that the 248-amino acid SFTPA2 protein contains a collagen-like region, a neck domain, and a C-terminal carbohydrate-recognition domain. SFTPA2 shares greater than 98% amino acid identity with SFTPA1 across the coding region and greater than 90% amino acid identity across noncoding regions.


Gene Structure

Katyal et al. (1992) determined that the SFTPA2 gene contains 5 exons.


Mapping

Kolble et al. (1993) presented evidence for a cluster of pulmonary surfactant-associated proteins on chromosome 10q22-q23. They constructed a phylogenetic tree showing the relationships of SPAI (SFTPA1), SPAII, mannose-binding lectin (MBL; 154545), SPB (178640), and SPC (178620) proteins.

Stumpf (2021) mapped the SFTPA2 gene to chromosome 10q22.3 based on an alignment of the SFTPA2 sequence (GenBank BC111571) with the genomic sequence (GRCh38).


Molecular Genetics

Surfactant protein-A, which plays a role in innate host defense in the lung, is also expressed in the eustachian tube. Ramet et al. (2001) reported that the frequency of specific surfactant protein-A haplotypes and genotypes differs between children with recurrent otitis media compared with a control population in Finland.

In affected members of 2 families with interstitial lung disease-2 (ILD2; 178500), some of whom also had lung cancer, Wang et al. (2009) identified 2 heterozygous missense mutations in the SFTPA2 gene (G231V, 178642.0001 and F198S, 178642.0002, respectively). Both mutations involve highly conserved residues in the carbohydrate recognition domain (CRD) and were predicted to disrupt protein structure; transfection studies in A549 human alveolar epithelial cells showed that the mutant proteins were retained in the endoplasmic reticulum and not secreted.

In 11 probands with ILD2, Legendre et al. (2020) identified heterozygous missense mutations in the SFTPA2 gene (see, e.g, 178642.0003-178642.0005). The mutations, which were found by direct sequencing, were either absent from or found at a low frequency in the gnomAD database. The mutation segregated with the disorder in families from whom DNA was available; there was evidence of incomplete penetrance and variable expressivity. In vitro functional expression studies in HEK293T cells transfected with the mutations showed normal protein expression with decreased secretion of SFTPA1 compared to controls. Abnormal cytoplasmic retention of mutant SFTPA1 in the alveolar epithelium was considered to contribute to pathogenicity.


ALLELIC VARIANTS 5 Selected Examples):

.0001   INTERSTITIAL LUNG DISEASE 2

SFTPA2, GLY231VAL
SNP: rs121917737, ClinVar: RCV000014086, RCV002509155

In affected members of a 4-generation family (F27) with interstitial lung disease-2 (ILD2; 178500), some of whom also had lung cancer, Wang et al. (2009) identified heterozygosity for a c.692G-T transversion in exon 6 of the SFTPA2 gene, resulting in a gly231-to-val (G231V) substitution at a highly conserved residue in the carbohydrate recognition domain (CRD). The mutation was not found in 3,557 population-based controls. Transfection studies in A549 human alveolar epithelial cells showed that the mutant protein was retained in the endoplasmic reticulum and not secreted.


.0002   INTERSTITIAL LUNG DISEASE 2

SFTPA2, PHE198SER
SNP: rs121917738, ClinVar: RCV000014087

In a 45-year-old Caucasian male (CKG810) with interstitial lung disease-2 (ILD2; 178500) and lung cancer (see 211980), Wang et al. (2009) identified heterozygosity for a c.593T-C transition in exon 6 of the SFTPA2 gene, resulting in a phe198-to-ser (F198S) substitution at a highly conserved residue in the carbohydrate recognition domain (CRD). The proband reported multiple family members with pulmonary fibrosis and/or lung cancer, but they were not available for study. The patient had no personal smoking history, but smoking status of his childhood home was unclear. The mutation was not found in 3,557 population-based controls. Transfection studies in A549 human alveolar epithelial cells showed that the mutant protein was retained in the endoplasmic reticulum and not secreted.


.0003   INTERSTITIAL LUNG DISEASE 2

SFTPA2, ASN171ILE
SNP: rs2132046635, ClinVar: RCV001780092

In a patient (family 4) with interstitial lung disease-2 (ILD2; 178500), Legendre et al. (2020) identified a heterozygous c.512A-T transversion (c.512A-T, NM_001098668.2) in exon 6 of the SFTPA2 gene, resulting in an asn171-to-ile (N171I) substitution in the CRD domain. The mutation, which was found by direct sequencing, was not found in the gnomAD database. In vitro functional expression studies in HEK293T cells transfected with the mutation showed normal protein expression with decreased secretion of SFTPA1 compared to controls. Abnormal cytoplasmic retention of mutant SFTPA1 in the alveolar epithelium was considered to contribute to pathogenicity. The patient had a significant family history of similar lung disease, but DNA from affected family members was not available for segregation studies.


.0004   INTERSTITIAL LUNG DISEASE 2

SFTPA2, VAL178MET ({dbSNP rs371035540})
SNP: rs371035540, gnomAD: rs371035540, ClinVar: RCV001265772, RCV001732099, RCV001780219

In 3 patients from 2 unrelated families (families 5 and 6) with interstitial lung disease-2 (ILD2; 178500), Legendre et al. (2020) identified a heterozygous c.532G-A transition (c.532G-A, NM_001098668.2) in exon 6 of the SFTPA2 gene, resulting in a val178-to-met (V178M) substitution in the CRD domain. The mutation, which was found by direct sequencing, was found at a low frequency in the gnomAD database (3 of 250,974 alleles). One unaffected family member in family 5 carried the mutation, consistent with incomplete penetrance. In vitro functional expression studies in HEK293T cells transfected with the mutation showed normal protein expression with decreased secretion of SFTPA1 compared to controls. Abnormal cytoplasmic retention of mutant SFTPA1 in the alveolar epithelium was considered to contribute to pathogenicity. The proband in family 6 had 2 similarly affected deceased family members, including those with lung cancer, but DNA from those individuals was not available for segregation studies.


.0005   INTERSTITIAL LUNG DISEASE 2

SFTPA2, TRP233ARG
SNP: rs2132045183, ClinVar: RCV001784080

In a mother and son (family 8) with interstitial lung disease-2 (ILD2; 178500), Legendre et al. (2020) identified a heterozygous c.697T-A transversion (c.697T-A, NM_001098668.2) in exon 6 of the SFTPA2 gene, resulting in a trp233-to-arg (W233R) substitution in the CRD domain. The mutation, which was found by direct sequencing, was not present in the gnomAD database. In vitro functional expression studies in HEK293T cells transfected with the mutation showed normal protein expression with decreased secretion of SFTPA2 compared to controls. Abnormal cytoplasmic retention of mutant SFTPA2 in the alveolar epithelium was considered to contribute to pathogenicity. There was a significant family history of similar lung disease, including lung cancer, but DNA from affected family members was not available for segregation studies.


REFERENCES

  1. Katyal, S. L., Singh, G., Locker, J. Characterization of a second human pulmonary surfactant-associated protein SP-A gene. Am. J. Resp. Cell Molec. Biol. 6: 446-452, 1992. [PubMed: 1372511] [Full Text: https://doi.org/10.1165/ajrcmb/6.4.446]

  2. Kolble, K., Lu, J., Mole, S. E., Kaluz, S., Reid, K. B. M. Assignment of the human pulmonary surfactant protein D gene (SFTP4) to 10q22-q23 close to the surfactant protein A gene cluster. Genomics 17: 294-298, 1993. [PubMed: 8406480] [Full Text: https://doi.org/10.1006/geno.1993.1324]

  3. Legendre, M., Butt, A., Borie, R., Debray, MP., Bouvry, D., Filhol-Blin, E., Desroziers, T., Nau, V., Copin, B., Dastot-Le Moal, F., Hery, M., Duquesnoy, P., and 29 others. Functional assessment and phenotypic heterogeneity of SFTPA1 and SFTPA2 mutations in interstitial lung diseases and lung cancer. Europ. Resp. J. 56: 2002806, 2020. [PubMed: 32855221] [Full Text: https://doi.org/10.1183/13993003.02806-2020]

  4. Ramet, M., Lofgren, J., Albo, O.-P., Hallman, M. Surfactant protein-A gene locus associated with recurrent otitis media. J. Pediat. 138: 266-268, 2001. [PubMed: 11174628] [Full Text: https://doi.org/10.1067/mpd.2001.110133]

  5. Stumpf, A. M. Personal Communication. Baltimore, Md. 11/16/2021.

  6. Wang, Y., Kuan, P. J., Xing, C., Cronkhite, J. T., Torres, F., Rosenblatt, R. L., DiMaio, J. M., Kinch, L. N., Grishin, N. V., Garcia, C. K. Genetic defects in surfactant protein A2 are associated with pulmonary fibrosis and lung cancer. Am. J. Hum. Genet. 84: 52-59, 2009. [PubMed: 19100526] [Full Text: https://doi.org/10.1016/j.ajhg.2008.11.010]


Contributors:
Anne M. Stumpf - updated : 11/18/2021
Cassandra L. Kniffin - updated : 11/15/2021
Marla J. F. O'Neill - updated : 2/19/2009
Ada Hamosh - updated : 4/23/2001

Creation Date:
Victor A. McKusick : 9/27/1993

Edit History:
carol : 11/19/2021
alopez : 11/18/2021
ckniffin : 11/15/2021
wwang : 02/27/2009
terry : 2/19/2009
mgross : 3/10/2003
cwells : 5/9/2001
terry : 4/23/2001
mcapotos : 6/8/2000
mcapotos : 6/7/2000
carol : 6/19/1998
terry : 6/18/1998
mimadm : 4/13/1994
carol : 9/27/1993